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Three homeodomain proteins define dorsoventral columns within the Drosophila central nervous system. Shown here is an immunohistochemical analysis of the neuroectoderm of stage 9 Drosophila embryos. Embryos were stained with antibodies to Vnd (ventral; pink-red), Ind (intermediate; brown), and Msh (dorsal; purple-blue). Anterior is up. The background is an enlargement of the inset image. (For details, see Weiss et al., p. 3591. See also McDonald et al., p. 3603, and Chu et al., p. 3613.)