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Alterations in actin cytoskeleton organization during abscisic acid-induced stomatal pore closure in Arabidopsis thaliana. Shown here are wild-type guard cells from a transgenic line of Arabidopsis expressing green fluorescent protein (GFP-mTn) targeted to the actin cytoskeleton. In the absence of abscisic acid (ABA), the guard cell remains open and the actin cables are transversely oriented (left). In the presence of ABA, actin cables become disorganized and shortened (right). Inactivation of the small GTPase AtRac1 is a limiting step in the ABA-induced stomatal closure process. (For details, see Lemichez et al., p. 1808.)